On the 300C a high pressure area or (stagnation point) is formed at the point of impact. The point of impact is within the front grill area on the 300C. For proper cooling it should be close to the center of the grill. Those "large" vertical and horizontal slats on the stock grill are there for a protective reasons yet they have a minimal surface area. Using a tight mesh grill seems like it would work as well but it usually it won't. The tighter the mesh the less air that will enter. At the point of impact the air flow is head on. As you move up down or left or right the air flow is angular. If you look at a tight mesh grill from an angle you'll notice that light will be blocked as the angle increases. Air will also tend to roll around a tight mesh grill at these angles.
There are other variables like frontal area, drag coefficient, air velocity, angle of attack, turbulent flow, transitional flow, laminar flow, wake vortices, boundary layer, skin-friction, eddies and air turbulence.
It would be nice if someone designed a aftermarket grill with both increased airflow and better looks in mind. Where the grill was actually flow tested in a wind tunnel.